Acoustic deterrent devices (ADDs) are used to deter seals from aquacultures but exposure of harbour porpoises (Phocoena phocoena) occurs as a side-effect. At construction sites, by contrast, ADDs are used to deter harbour porpoises from the zone in which pile driving noise can induce temporary threshold shifts (TTSs). ADDs emit such high pressure levels that there is concern that ADDs themselves may induce a TTS. A harbour porpoise in human care was exposed to an artificial ADD signal with a peak frequency of 14 kHz. A significant TTS was found, measured by auditory evoked potentials, with an onset of 142 dB re 1 μPa2s at 20 kHz and 147 dB re 1 μPa2s at 28 kHz. The authors therefore strongly recommend to gradually increase and down regulate source levels of ADDs to the desired deterrence range. However, further research is needed to develop a reliable relationship between received levels and deterrence.

1.
Ainslie
,
M. A.
, and
McColm
,
J. G.
(
1998
). “
A simplified formula for viscous and chemical absorption in sea water
,”
J. Acoust. Soc. Am.
103
,
1671
1672
.
2.
Andreasen
,
H.
,
Ross
,
S. D.
,
Siebert
,
U.
,
Andersen
,
N. G.
,
Ronnenberg
,
K.
, and
Gilles
,
A.
(
2017
). “
Diet composition and food consumption rate of harbor porpoises (Phocoena phocoena) in the western Baltic Sea
,”
Mar. Mammal Sci.
33
,
1053
1079
.
3.
ASCOBANS
(
2002
). “
Recovery plan for Baltic harbour porpoises (Jastarnia Plan)
.”
4.
ASCOBANS
(
2012
). Conservation of Harbour Porpoises and Adoption of a Conservation Plan for the Western Baltic, the Belt Sea and Kattegat Brighton, United Kingdom, October 22–24, 2012,
46
pp.
5.
Bailey
,
H.
,
Senior
,
B.
,
Simmons
,
D.
,
Rusin
,
J.
,
Picken
,
G.
, and
Thompson
,
P. M.
(
2010
). “
Assessing underwater noise levels during pile-driving at an offshore windfarm and its potential effects on marine mammals
,”
Mar. Pollut. Bull.
60
,
888
897
.
6.
Beineke
,
A.
,
Siebert
,
U.
,
McLachlan
,
M.
,
Bruhn
,
R.
,
Thron
,
K.
,
Failing
,
K.
,
Müller
,
G.
, and
Baumgärtner
,
W.
(
2005
). “
Investigations of the potential influence of environmental contaminants on the thymus and spleen of harbor porpoises (Phocoena phocoena)
,”
Environ. Sci. Technol.
39
,
3933
3938
.
7.
BMU
(
2014
).
Concept for the Protection of Harbour Porpoises from Sound Exposures during the Construction of Offshore Wind Farms in the German North Sea (Sound Protection Concept),
21st ASCOBANS Advisory Committee Meeting AC21/Inf.3.2.2.a (P) Gothenburg, Sweden (September 29–October 1, 2014),
33
pp.
8.
Brandt
,
M. J.
,
Dragon
,
A. C.
,
Diederichs
,
A.
,
Bellmann
,
M. A.
,
Wahl
,
V.
,
Piper
,
W.
,
Nabe-Nielsen
,
J.
, and
Nehls
,
G.
(
2018
). “
Disturbance of harbour porpoises during construction of the first seven offshore wind farms in Germany
,”
Mar. Ecol. Prog. Ser.
596
,
213
232
.
9.
Brandt
,
M. J.
,
Höschle
,
C.
,
Diederichs
,
A.
,
Betke
,
K.
,
Matuschek
,
R.
, and
Nehls
,
G.
(
2013
). “
Seal scarers as a tool to deter harbour porpoises from offshore construction sites
,”
Mar. Ecol. Prog. Ser.
475
,
291
302
.
10.
Brandt
,
M. J.
,
Höschle
,
C.
,
Diederichs
,
A.
,
Betke
,
K.
,
Matuschek
,
R.
,
Witte
,
S.
, and
Nehls
,
G.
(
2012
). “
Far-reaching effects of a seal scarer on harbour porpoises, Phocoena phocoena
,”
Aquat. Conserv. Mar. Freshw. Ecosyst.
23
,
222
232
.
11.
Clausen
,
K. T.
,
Wahlberg
,
M.
,
Beedholm
,
K.
,
DeRuiter
,
S.
, and
Madsen
,
P. T.
(
2010
). “
Click communication in harbour porpoises Phocoena phocoena
,”
Bioacoust.
20
,
1
28
.
12.
Coram
,
A.
,
Gordon
,
J.
,
Thompson
,
D.
, and
Northridge
,
S.
(
2014
). Evaluating and assessing the relative effectiveness of Acoustic Deterrent Devices and other non-lethal measures on marine mammals, Scottish Government, St Andrews, Scotland.
13.
Dähne
,
M.
,
Gilles
,
A.
,
Lucke
,
K.
,
Peschko
,
V.
,
Adler
,
S.
,
Krügel
,
K.
,
Sundermeyer
,
J.
, and
Siebert
,
U.
(
2013
). “
Effects of pile-driving on harbour porpoises (Phocoena phocoena) at the first offshore wind farm in Germany
,”
Environ. Res. Lett.
8
,
025002
.
14.
Dähne
,
M.
,
Tougaard
,
J.
,
Carstensen
,
J.
,
Rose
,
A.
, and
Nabe-Nielsen
,
J.
(
2017
). “
Bubble curtains attenuate noise from offshore wind farm construction and reduce temporary habitat loss for harbour porpoises
,”
Mar. Ecol. Prog. Ser.
580
,
221
237
.
15.
Das
,
K.
,
Vossen
,
A.
,
Tolley
,
K.
,
Víkingsson
,
G.
,
Thron
,
K.
,
Müller
,
G.
,
Baumgärtner
,
W.
, and
Siebert
,
U.
(
2006
). “
Interfollicular fibrosis in the thyroid of the harbour porpoise: An endocrine disruption?
,”
Arch. Environ. Contam. Toxicol.
51
,
720
729
.
16.
DeRuiter
,
S. L.
,
Bahr
,
A.
,
Blanchet
,
M.-A.
,
Hansen
,
S. F.
,
Kristensen
,
J. H.
,
Madsen
,
P. T.
,
Tyack
,
P. L.
, and
Wahlberg
,
M.
(
2009
). “
Acoustic behaviour of echolocating porpoises during prey capture
,”
J. Exp. Biol.
212
,
3100
3107
.
17.
Don
,
M.
,
Elberling
,
C.
, and
Waring
,
M.
(
1984
). “
Objective detection of averaged auditory brainstem responses
,”
Int. J. Audiol.
13
,
219
228
.
18.
Dyndo
,
M.
,
Wiśniewska
,
D. M.
,
Rojano-Doñate
,
L.
, and
Madsen
,
P. T.
(
2015
). “
Harbour porpoises react to low levels of high frequency vessel noise
,”
Sci. Rep.
5
,
11083
.
19.
Elberling
,
C.
, and
Don
,
M.
(
1984
). “
Quality estimation of averaged auditory brainstem responses
,”
Scand. Audiol.
13
,
187
197
.
20.
FAO
(
2018
).
The State of World Fisheries and Aquacultures. Meeting the Sustainable Development Goals
, Rome, Italy, License: CC BY-NC-SA 3.0 IGO, pp.
1
227
.
21.
Findlay
,
C. R.
,
Ripple
,
H. D.
,
Coomber
,
F.
,
Froud
,
K.
,
Harries
,
O.
,
van Geel
,
N. C. F.
,
Calderan
,
S. V.
,
Benjamins
,
S.
,
Risch
,
D.
, and
Wilson
,
B.
(
2018
). “
Mapping widespread and increasing underwater noise pollution from acoustic deterrent devices
,”
Mar. Pollut. Bull.
135
,
1042
1050
.
22.
Finneran
,
J. J.
(
2009
). “
Evoked response study tool: A portable, rugged system for single and multiple auditory evoked potential measurements
,”
J. Acoust. Soc. Am.
126
,
491
500
.
23.
Finneran
,
J. J.
(
2018
). “
Conditioned attenuation of auditory brainstem responses in dolphins warned of an intense noise exposure: Temporal and spectral patterns
,”
J. Acoust. Soc. Am.
143
,
795
810
.
24.
Finneran
,
J. J.
,
Carder
,
D. A.
,
Schlundt
,
C. E.
, and
Dear
,
R. L.
(
2010a
). “
Growth and recovery of temporary threshold shift at 3 kHz in bottlenose dolphins: Experimental data and mathematical models
,”
J. Acoust. Soc. Am.
127
,
3256
3266
.
25.
Finneran
,
J. J.
,
Carder
,
D. A.
,
Schlundt
,
C. E.
, and
Dear
,
R. L.
(
2010b
). “
Temporary threshold shift in a bottlenose dolphin (Tursiops truncatus) exposed to intermittent tones
,”
J. Acoust. Soc. Am.
127
,
3267
3272
.
26.
Finneran
,
J. J.
,
Carder
,
D. A.
,
Schlundt
,
C. E.
, and
Ridgway
,
S. H.
(
2005
). “
Temporary threshold shift in bottlenose dolphins (Tursiops truncatus) exposed to mid-frequency tones
,”
J. Acoust. Soc. Am.
118
,
2696
2705
.
27.
Finneran
,
J. J.
,
Houser
,
D. S.
,
Blasko
,
D.
,
Hicks
,
C.
,
Hudson
,
J.
, and
Osborn
,
M.
(
2008
). “
Estimating bottlenose dolphin (Tursiops truncatus) hearing thresholds from single and multiple simultaneous auditory evoked potentials
,”
J. Acoust. Soc. Am.
123
,
542
551
.
28.
Gilles
,
A.
,
Viquerat
,
S.
,
Becker
,
E. A.
,
Forney
,
K. A.
,
Geelhoed
,
S. C. V.
,
Haelters
,
J.
,
Nabe-Nielsen
,
J.
,
Scheidat
,
M.
,
Siebert
,
U.
,
Sveegaard
,
S.
,
van Beest
,
F. M.
,
van Bemmelen
,
R.
, and
Aarts
,
G.
(
2016
). “
Seasonal habitat-based density models for a marine top predator, the harbor porpoise, in a dynamic environment
,”
Ecosphere
7
,
e01367
.
29.
Götz
,
T.
, and
Janik
,
V.
(
2013
). “
Acoustic deterrent devices to prevent pinniped depredation: Efficiency, conservation concerns and possible solutions
,”
Mar. Ecol. Prog. Ser.
492
,
285
302
.
30.
Hammond
,
P. S.
,
Lacey
,
C.
,
Gilles
,
A.
,
Viquerat
,
S.
,
Börjesson
,
P.
,
Herr
,
H.
,
Macleod
,
K.
,
Ridoux
,
V.
,
Santos
,
M. B.
,
Scheidat
,
M.
,
Teilmann
,
J.
,
Vingada
,
J.
, and
Øien
,
N.
(
2017
). Estimates of Cetacean Abundance in European Atlantic Waters in Summer 2016 from the SCANS-III Aerial and Shipboard Surveys, pp.
1
39
.
32.
Jepson
,
P. D.
,
Deaville
,
R.
,
Barber
,
J. L.
,
Aguilar
,
À.
,
Borrell
,
A.
,
Murphy
,
S.
,
Barry
,
J.
,
Brownlow
,
A.
,
Barnett
,
J.
,
Berrow
,
S.
,
Cunningham
,
A. A.
,
Davison
,
N. J.
,
ten Doeschate
,
M.
,
Esteban
,
R.
,
Ferreira
,
M.
,
Foote
,
A. D.
,
Genov
,
T.
,
Giménez
,
J.
,
Loveridge
,
J.
,
Llavona
,
Á.
,
Martin
,
V.
,
Maxwell
,
D. L.
,
Papachlimitzou
,
A.
,
Penrose
,
R.
,
Perkins
,
M. W.
,
Smith
,
B.
,
de Stephanis
,
R.
,
Tregenza
,
N.
,
Verborgh
,
P.
,
Fernandez
,
A.
, and
Law
,
R. J.
(
2016
). “
PCB pollution continues to impact populations of orcas and other dolphins in European waters
,”
Sci. Rep.
6
,
18573
.
33.
Kastak
,
D.
, and
Schusterman
,
R. J.
(
1998
). “
Low-frequency amphibious hearing in pinnipeds: Methods, measurements, noise, and ecology
,”
J. Acoust. Soc. Am.
103
,
2216
2228
.
34.
Kastelein
,
R. A.
,
Bunskoek
,
P.
,
Hagedoorn
,
M.
,
Au
,
W. W. L.
, and
de Haan
,
D.
(
2002
). “
Audiogram of a harbor porpoise (Phocoena phocoena) measured with narrow-band frequency-modulated signals
,”
J. Acoust. Soc. Am.
112
,
334
344
.
35.
Kastelein
,
R. A.
,
Hoek
,
L.
,
de Jong
,
C. A. F.
, and
Wensveen
,
P. J.
(
2010
). “
The effect of signal duration on the underwater detection thresholds of a harbor porpoise (Phocoena phocoena) for single frequency-modulated tonal signals between 0.25 and 160 kHz
,”
J. Acoust. Soc. Am.
128
,
3211
3222
.
36.
Kastelein
,
R. A.
,
Gransier
,
R.
,
Marijt
,
M. A. T.
, and
Hoek
,
L.
(
2015a
). “
Hearing frequency thresholds of harbor porpoises (Phocoena phocoena) temporarily affected by played back offshore pile driving sounds
,”
J. Acoust. Soc. Am.
137
,
556
564
.
37.
Kastelein
,
R. A.
,
Gransier
,
R.
,
Schop
,
J.
, and
Hoek
,
L.
(
2015b
). “
Effects of exposure to intermittent and continuous 6-7 kHz sonar sweeps on harbor porpoise (Phocoena phocoena) hearing
,”
J. Acoust. Soc. Am.
137
,
1623
1633
.
38.
Kastelein
,
R. A.
,
Helder-Hoek
,
L.
,
Covi
,
J.
, and
Gransier
,
R.
(
2016
). “
Pile driving playback sounds and temporary threshold shift in harbor porpoises (Phocoena phocoena): Effect of exposure duration
,”
J. Acoust. Soc. Am.
139
,
2842
2851
.
39.
Kastelein
,
R. A.
,
Helder-Hoek
,
L.
, and
Terhune
,
J. M.
(
2018
). “
Hearing thresholds, for underwater sounds, of harbor seals (Phoca vitulina) at the water surface
,”
J. Acoust. Soc. Am.
143
,
2554
2563
.
40.
Kastelein
,
R. A.
,
Hoek
,
L.
,
Gransier
,
R.
,
Rambags
,
M.
, and
Claeys
,
N.
(
2014a
). “
Effect of level, duration, and inter-pulse interval of 1-2 kHz sonar signal exposures on harbor porpoise hearing
,”
J. Acoust. Soc. Am.
136
,
412
422
.
41.
Kastelein
,
R. A.
,
Schop
,
J.
,
Gransier
,
R.
, and
Hoek
,
L.
(
2014b
). “
Frequency of greatest temporary hearing threshold shift in harbor porpoises (Phocoena phocoena) depends on the noise level
,”
J. Acoust. Soc. Am.
136
,
1410
1418
.
42.
Kastelein
,
R. A.
,
Wensveen
,
P. J.
,
Hoek
,
L.
,
Verboom
,
W. C.
, and
Terhune
,
J. M.
(
2009
). “
Underwater detection of tonal signals between 0.125 and 100 kHz by harbor seals (Phoca vitulina)
,”
J. Acoust. Soc. Am.
125
,
1222
1229
.
43.
King
,
S. L.
,
Schick
,
R. S.
,
Donovan
,
C.
,
Booth
,
C. G.
,
Burgman
,
M.
,
Thomas
,
L.
, and
Harwood
,
J.
(
2015
). “
An interim framework for assessing the population consequences of disturbance
,”
Methods Ecol. Evol.
6
,
1150
1158
.
44.
Koch
,
M.
, and
Schnitzler
,
H.-U.
(
1997
). “
The acoustic startle response in rats—circuits mediating evocation, inhibition and potentiation
,”
Behav. Brain Res.
89
,
35
49
.
45.
Lepper
,
P. A.
,
Turner
,
V. L. G.
,
Goodson
,
A. D.
, and
Black
,
K. D.
(
2004
). “
Source levels and spectra emitted by three commercial aquaculture anti-predation devices
,” in
Proceedings of the 7th European Conference on Underwater Acoustics
(July 5–8, 2004), Vol.
6
,
6
pp.
46.
Lucke
,
K.
,
Siebert
,
U.
,
Lepper
,
P. A.
, and
Blanchet
,
M.-A.
(
2009
). “
Temporary shift in masked hearing thresholds in a harbor porpoise (Phocoena phocoena) after exposure to seismic airgun stimuli
,”
J. Acoust. Soc. Am.
125
,
4060
4070
.
47.
Madsen
,
P. T.
(
2005
). “
Marine mammals and noise: Problems with root mean square sound pressure levels for transients
,”
J. Acoust. Soc. Am.
117
,
3952
3957
.
48.
Mahfouz
,
C.
,
Henry
,
F.
,
Courcot
,
L.
,
Pezeril
,
S.
,
Bouveroux
,
T.
,
Dabin
,
W.
,
Jauniaux
,
T.
,
Khalaf
,
G.
, and
Amara
,
R.
(
2014
). “
Harbour porpoises (Phocoena phocoena) stranded along the southern North Sea: An assessment through metallic contamination
,”
Environ. Res.
133
,
266
273
.
49.
Mann
,
D.
,
Hill-Cook
,
M.
,
Manire
,
C.
,
Greenhow
,
D.
,
Montie
,
E.
,
Powell
,
J.
,
Wells
,
R.
,
Bauer
,
G.
,
Cunningham-Smith
,
P.
,
Lingenfelser
,
R.
,
DiGiovanni
,
R.
,
Stone
,
A.
,
Brodsky
,
M.
,
Stevens
,
R.
,
Kieffer
,
G.
, and
Hoetjes
,
P.
(
2010
). “
Hearing loss in stranded odontocete dolphins and whales
,”
PLoS One
5
,
e13824
.
50.
McFadden
,
D.
, and
Plattsmier
,
H. S.
(
1983
). “
Frequency patterns of TTS for different exposure intensities
,”
J. Acoust. Soc. Am.
74
,
1178
1184
.
51.
Mikkelsen
,
L.
,
Hermannsen
,
L.
,
Beedholm
,
K.
,
Madsen
,
P. T.
, and
Tougaard
,
J.
(
2017
). “
Simulated seal scarer sounds scare porpoises, but not seals: Species-specific responses to 12 kHz deterrence sounds
,”
R. Soc. Open Sci.
4
,
170286
.
52.
Mooney
,
T. A.
,
Nachtigall
,
P. E.
,
Breese
,
M.
,
Vlachos
,
S.
, and
Au
,
W. W. L.
(
2009
). “
Predicting temporary threshold shifts in a bottlenose dolphin (Tursiops truncatus): The effects of noise level and duration
,”
J. Acoust. Soc. Am.
125
,
1816
1826
.
53.
Morell
,
M.
,
Brownlow
,
A.
,
McGovern
,
B.
,
Raverty
,
S. A.
,
Shadwick
,
R. E.
, and
André
,
M.
(
2017
). “
Implementation of a method to visualize noise-induced hearing loss in mass stranded cetaceans
,”
Sci. Rep.
7
,
41848
.
54.
Mulsow
,
J.
, and
Reichmuth
,
C.
(
2010
). “
Psychophysical and electrophysiological aerial audiograms of a Steller sea lion (Eumetopias jubatus)
,”
J. Acoust. Soc. Am.
127
,
2692
2701
.
55.
Nachtigall
,
P. E.
,
Supin
,
A. Y.
,
Pacini
,
A. F.
, and
Kastelein
,
R. A.
(
2016
). “
Conditioned hearing sensitivity change in the harbor porpoise (Phocoena phocoena)
,”
J. Acoust. Soc. Am.
140
,
960
967
.
56.
Nachtigall
,
P. E.
,
Supin
,
A. Y.
,
Pacini
,
A. F.
, and
Kastelein
,
R. A.
(
2017
). “
Four odontocete species change hearing levels when warned of impending loud sound
,”
Integr. Zool.
13
,
160
165
57.
National Marine Fisheries Service
(
2018
). 2018 Revisions to: “
Technical guidance for assessing the effects of anthropogenic sound on marine mammal hearing (Version 2.0): Underwater thresholds for onset of permanent and temporary threshold shifts
,”
167
pp.
58.
National Oceanic and Atmospheric Administration; National Marine Fisheries Service
(
2016
). “
Technical guidance for assessing the effects of anthropogenic sound on marine mammal hearing: Underwater acoustic thresholds for onset of permanent and temporary threshold shifts
,” Silver Spring, MD,
178
pp.
59.
Olesiuk
,
P. F.
,
Nichol
,
L. M.
,
Sowden
,
M. J.
, and
Ford
,
J. K. B.
(
2002
). “
Effect of the sound generated by an acoustic harassment device on the relative abundance and distribution of harbor porpoises (Phocoena phocoena) in Retreat Passage, British Columbia
,”
Mar. Mammal Sci.
18
,
843
862
.
60.
Peschko
,
V.
,
Ronnenberg
,
K.
,
Siebert
,
U.
, and
Gilles
,
A.
(
2016
). “
Trends of harbour porpoise (Phocoena phocoena) density in the southern North Sea
,”
Ecol. Indic.
60
,
174
183
.
61.
Pilz
,
P. K. D.
,
Schnitzler
,
H.-U.
, and
Menne
,
D.
(
1987
). “
Acoustic startle threshold of the albino rat (Rattus norvegicus)
,”
J. Comp. Psychol.
101
,
67
72
.
62.
Popov
,
V. V.
,
Supin
,
A. Y.
,
Rozhnov
,
V. V.
,
Nechaev
,
D. I.
, and
Sysueva
,
E. V.
(
2014
). “
The limits of applicability of the sound exposure level (SEL) metric to temporal threshold shifts (TTS) in beluga whales, Delphinapterus leucas
,”
J. Exp. Biol.
217
,
1804
1810
.
63.
Popov
,
V. V.
,
Supin
A. Y.
,
Wang
,
D.
,
Wang
,
K.
,
Dong
,
L.
, and
Wang
,
S.
(
2011
). “
Noise-induced temporary threshold shift and recovery in Yangtze finless porpoises Neophocaena phocaenoides asiaeorientalis
,”
J. Acoust. Soc. Am.
130
,
574
584
.
64.
R Core Team
(
2019
). “
R: A language and environment for statistical computing
,” R Foundation for Statistical Computing, Vienna, Austria. https://www.R-project.org/ (Last viewed November 25, 2019).
65.
Reeves
,
R. R.
,
McClellan
,
K.
, and
Werner
,
T. B.
(
2013
). “
Marine mammal bycatch in gillnet and other entangling net fisheries, 1990 to 2011
,”
Endanger. Species Res.
20
,
71
97
.
66.
Reichmuth
,
C.
,
Holt
,
M. M.
,
Mulsow
,
J.
,
Sills
,
J. M.
, and
Southall
,
B. L.
(
2013
). “
Comparative assessment of amphibious hearing in pinnipeds
,”
J. Comp. Physiol. A
199
,
491
507
.
67.
Remy
,
T.
, and
Mbistrova
,
A.
(
2018
). Offshore Wind in Europe—Key Trends and Statistics 2017, Brussels, pp.
1
37
.
68.
Ruser
,
A.
,
Dähne
,
M.
,
van Neer
,
A.
,
Lucke
,
K.
,
Sundermeyer
,
J.
,
Siebert
,
U.
,
Houser
,
D. S.
,
Finneran
,
J. J.
,
Everaarts
,
E.
,
Meerbeek
,
J.
,
Dietz
,
R.
,
Sveegaard
,
S.
, and
Teilmann
,
J.
(
2016
). “
Assessing auditory evoked potentials of wild harbor porpoises (Phocoena phocoena)
,”
J. Acoust. Soc. Am.
140
,
442
452
.
69.
Schlundt
,
C. E.
,
Finneran
,
J. J.
,
Carder
,
D. A.
, and
Ridgway
,
S. H.
(
2000
). “
Temporary shift in masked hearing thresholds of bottlenose dolphins, Tursiops truncatus, and white whales, Delphinapterus leucas, after exposure to intense tones
,”
J. Acoust. Soc. Am.
107
,
3496
3508
.
70.
Serpanos
,
Y. C.
,
O'Malley
,
H.
, and
Gravel
,
J. S.
(
1997
). “
The relationship between loudness intensity functions and the click-ABR wave V latency
,”
Ear Hear.
18
,
409
419
.
71.
Sørensen
,
P. M.
,
Wisniewska
,
D. M.
,
Jensen
,
F. H.
,
Johnson
,
M.
,
Teilmann
,
J.
, and
Madsen
,
P. T.
(
2018
). “
Click communication in wild harbour porpoises (Phocoena phocoena)
,”
Sci. Rep.
8
,
9702
.
72.
Southall
,
B. L.
,
Bowles
,
A. E.
,
Ellison
,
W. T.
,
Finneran
,
J. J.
,
Gentry
,
R. L.
,
Greene
,
C. R.
, Jr.
,
Kastak
,
D.
,
Ketten
,
D. R.
,
Miller
,
J. H.
,
Nachtigall
,
P. E.
,
Richardson
,
W. J.
,
Thomas
,
J. A.
, and
Tyack
,
P. L.
(
2007
). “
Marine mammal noise exposure criteria: Initial scientific recommendations
,”
Aquat. Mamm.
33
,
411
522
.
73.
Southall
,
B. L.
,
Finneran
,
J. J.
,
Reichmuth
,
C.
,
Nachtigall
,
P. E.
,
Ketten
,
D. R.
,
Bowles
,
A. E.
,
Ellison
,
W. T.
,
Nowacek
,
D. P.
, and
Tyack
,
P. L.
(
2019
). “
Marine mammal noise exposure criteria: Updated scientific recommendations for residual hearing effects
,”
Aquat. Mamm.
45
,
125
232
.
74.
Stöber
,
U.
, and
Thomsen
,
F.
(
2019
). “
Effect of impact pile driving noise on marine mammals: A comparison of different noise exposure criteria
,”
J. Acoust. Soc. Am.
145
,
3252
3259
.
75.
Sueur
,
J.
,
Aubin
,
T.
, and
Simonis
,
C.
(
2008
). “
Seewave: A free modular tool for sound analysis and synthesis
,”
Bioacoustics
18
,
213
226
.
76.
Tougaard
,
J.
, and
Beedholm
,
K.
(
2018
). “
Practical implementation of auditory time and frequency weighting in marine bioacoustics
,”
Appl. Acoust.
145
,
137
143
.
77.
Tougaard
,
J.
,
Carstensen
,
J.
,
Teilmann
,
J.
,
Skov
,
H.
, and
Rasmussen
,
P.
(
2009
). “
Pile driving zone of responsiveness extends beyond 20 km for harbor porpoises (Phocoena phocoena (L.))
,”
J. Acoust. Soc. Am.
126
,
11
14
.
78.
Tougaard
,
J.
,
Wright
,
A. J.
, and
Madsen
,
P. T.
(
2015
). “
Cetacean noise criteria revisited in the light of proposed exposure limits for harbour porpoises
,”
Mar. Pollut. Bull.
90
,
196
208
.
79.
Villadsgaard
,
A.
,
Wahlberg
,
M.
, and
Tougaard
,
J.
(
2007
). “
Echolocation signals of wild harbour porpoises, Phocoena phocoena
,”
J. Exp. Biol.
210
,
56
64
.
80.
Viquerat
,
S.
,
Herr
,
H.
,
Gilles
,
A.
,
Peschko
,
V.
,
Siebert
,
U.
,
Sveegaard
,
S.
, and
Teilmann
,
J.
(
2014
). “
Abundance of harbour porpoises (Phocoena phocoena) in the western Baltic, Belt Seas and Kattegat
,”
Mar. Biol.
161
,
745
754
.
81.
Wisniewska
,
D. M.
,
Johnson
,
M.
,
Teilmann
,
J.
,
Siebert
,
U.
,
Galatius
,
A.
,
Dietz
,
R.
, and
Madsen
,
P. T.
(
2018
). “
High rates of vessel noise disrupt foraging in wild harbour porpoises (Phocoena phocoena)
,”
Proc. R. Soc. B Biol. Sci.
285
,
2017
2314
.
82.
Wisniewska
,
D. M. M.
,
Johnson
,
M.
,
Teilmann
,
J.
,
Rojano-Doñate
,
L.
,
Shearer
,
J.
,
Sveegaard
,
S.
,
Miller
,
L. A. A.
,
Siebert
,
U.
, and
Madsen
,
P. T. T.
(
2016
). “
Ultra-high foraging rates of harbor porpoises make them vulnerable to anthropogenic disturbance
,”
Curr. Biol.
26
,
1441
1446
.
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