It is difficult and expensive to match the sensitivity of the most sensitive vertebrate ears with off-the-shelf microphones due to the self-noise of the sensor. The extremely small apertures of microelectromechanical microphones create options to use horn waveguides to amplify sound prior to transduction without resulting in an unacceptably narrow directivity. Substantial gain can be achieved at wavelengths larger than the horn. An analytical model of an exponential horn embedded in a rigid spherical housing was formulated to describe the gain relative to a free-field receiver as a function of frequency and angle of arrival. For waves incident on-axis, the analytical model provided an accurate estimate of gain at high frequencies as validated by experimental measurement. Numerical models, using the equivalent source method, can account for higher order modes and comprehensively describe the acoustic scattering within and around the horn for waves arriving from any direction. Results show the directivity of horn receivers were adequately described by the analytical model up to a critical wavelength, and the mechanisms of deviation in gain at high frequencies and large angles of arrival were identified.

1.
D. T.
Blumstein
,
D. J.
Mennill
,
P.
Clemins
,
L.
Girod
,
K.
Yao
,
G.
Patricelli
,
J. L.
Deppe
,
A. H.
Krakauer
,
C.
Clark
,
K. A.
Cortopassi
, and
S. F.
Hanser
, “
Acoustic monitoring in terrestrial environments using microphone arrays: Applications, technological considerations, and prospectus
,”
J. Appl. Ecol.
48
,
758
767
(
2011
).
2.
M.
Campos Cerqueira
and
M. T.
Aide
, “
Improving distribution data of threatened species by combining acoustic monitoring and occupancy modeling
,”
Methods Ecol. Evol.
7
,
1340
1348
(
2016
).
3.
M. E.
Thompson
,
S. J.
Schwager
, and
K. B.
Payne
, “
Heard but not seen: An acoustic survey of the African forest elephant population at Kakum Conservation Area, Ghana
,”
Afr. J. Ecol.
48
,
224
231
(
2010
).
4.
T. A.
Marques
,
L.
Thomas
,
S. W.
Martin
,
D. K.
Mellinger
,
J. A.
Ward
,
D. J.
Moretti
,
D.
Harris
, and
P. L.
Tyack
, “
Estimating animal population density using passive acoustics
,”
Biol. Rev. Cambridge Philos. Soc.
88
,
287
309
(
2013
).
5.
D. J.
Mennitt
and
K. M.
Fristrup
, “
Influence factors and spatiotemporal patterns of environmental sound levels in the contiguous United States
,”
Noise Control Eng. J.
64
,
342
353
(
2016
).
6.
E.
Lynch
,
D.
Joyce
, and
K.
Fristrup
, “
An assessment of noise audibility and sound levels in U.S. National Parks
,”
Landsc. Ecol.
26
,
1297
1309
(
2011
).
7.
L.
Fritschi
,
L.
Brown
,
R.
Kim
,
D.
Schwela
, and
S.
Kephalopolous
,
Burden of Disease from Environmental Noise: Quantification of Healthy Life Years lost in Europe
(
World Health Organization
,
Bonn
,
2011
).
8.
G.
Shannon
,
M. F.
McKenna
,
L. M.
Angeloni
,
K. R.
Crooks
,
K. M.
Fristrup
,
E.
Brown
,
K. A.
Warner
,
M. D.
Nelson
,
C.
White
,
J.
Briggs
, and
S.
McFarland
, “
A synthesis of two decades of research documenting the effects of noise on wildlife
,”
Biol. Rev.
91
,
982
1005
(
2015
).
9.
B. C.
Pijanowski
,
A.
Farina
,
S. H.
Gage
,
S. L.
Dumyahn
, and
B. L.
Krause
, “
What is soundscape ecology? An introduction and overview of an emerging new science
,”
Landsc. Ecol.
26
,
1213
1232
(
2011
).
10.
R.
Horonjeff
, “
Overcoming instrument noise when measuring in quiet environments
,” in
Proceedings of Inter-noise
(
2002
).
11.
D. J.
Mennitt
,
K.
Fristrup
, and
K.
Sherrill
, “
A geospatial model of ambient sound pressure levels in the contiguous United States
,”
J. Acoust. Soc. Am.
135
,
2746
2764
(
2014
).
12.
E.
Lynch
,
L. M.
Angeloni
,
K.
Fristrup
,
D.
Joyce
, and
G.
Wittemyer
, “
The use of on-animal acoustical recording devices for studying animal behavior
,”
J. Ecol. Evol.
3
,
2030
2037
(
2013
).
13.
H. R.
Goerlitz
,
S.
Greif
, and
B. M.
Siemers
, “
Cues for acoustic detection of prey: Insect rustling sounds and the influence of walking substrate
,”
J. Exp. Biol.
211
.
2799
2806
(
2008
).
14.
R.
Fay
,
Hearing in Vertebrates: A Psychophysics Databook
(
Hill-Fay Associates
,
Winnetka, IL
,
1988
).
15.
S.
Wahlström
, “
The parabolic reflector as an acoustical amplifier
,”
J. Aud. Eng. Soc.
33
,
418
429
(
1985
).
16.
N. H.
Fletcher
, “
Acoustic systems in biology: From insects to elephants
,”
Acoust. Austr.
33
,
83
88
(
2005
).
17.
H. F.
Olson
and
I.
Wolff
, “
Sound concentrator for microphones”
J. Acoust. Soc. Am.
1
,
410
417
(
1930
).
18.
D. M.
Donskoy
and
B. A.
Cray
, “
Acoustic particle velocity horns
,”
J. Acoust. Soc. Am.
131
,
3883
3890
(
2012
).
19.
K. M.
Fristrup
and
D.
Mennitt
, “
Bioacoustical monitoring in terrestrial environments
,”
Acoust. Today
8
,
16
24
(
2012
).
20.
E.
Eisner
, “
Complete solutions of the ‘Webster’ horn equation
,”
J. Acoust. Soc. Am.
41
,
1126
1146
(
1967
).
21.
K. R.
Holland
,
F. J.
Fahy
, and
C. L.
Morfey
, “
Prediction and measurement of the one-parameter behavior of horns
,”
J. Aud. Eng. Soc.
39
,
315
337
(
1991
).
22.
M. R.
Stinson
and
G. A.
Daigle
, “
Comparison of an analytic horn equation approach and a boundary element method for the calculation of sound fields in the human ear canal
,”
J. Acoust. Soc. Am.
118
,
2405
2411
(
2005
).
23.
D. J.
Mennitt
and
K.
Fristrup
, “
Gain and directivity of exponential horn receivers
,”
J. Acoust. Soc. Am.
140
,
3140
(
2016
).
24.
P. M.
Morse
,
Vibration and Sound
(
Acoustical Society of America
,
New York
,
1981
).
25.
T. F.
Johansen
, “
On the directivity of horn loudspeakers
,”
J. Aud. Eng. Soc.
42
,
1008
1019
(
1994
).
26.
H. F.
Olson
,
Elements of Acoustical Engineering
(
D. Van Nostrand Company
,
New York
,
1947
).
27.
L.
Beranek
and
T.
Mellow
,
Acoustics: Sound Fields and Transducers
(
Academic Press
,
New York
,
2012
).
28.
V.
Salmon
, “
Horns
,” in
Encyclopedia of Acoustics
, edited by
M. J.
Crocker
(
John Wiley
,
New York
,
1997
), Vol. 4, pp.
1925
1931
.
29.
W. M.
Hall
, “
Comments on the Theory of Horns
,”
J. Acoust. Soc. Am.
3
,
552
561
(
1932
).
30.
IEC Standard 60268-4:
Sound System Equipment Part 4: Microphones
(
International Electrotechnical Commission
,
Geneva, Switzerland
,
2004
).
31.
B.
Zhou
and
D.
Green
, “
Characterization of external ear impulse responses using Golay codes
,”
J. Acoust. Soc. Am.
92
,
1169
1171
(
1992
).
32.
G. H.
Koopmann
,
L.
Song
, and
J. B.
Fahnline
, “
A method for computing acoustic fields based on the principle of wave superposition
,”
J. Acoust. Soc. Am.
86
,
2433
2438
(
1989
).
33.
M.
Johnson
,
S.
Elliott
,
K.
Baek
, and
J.
Garcia-Bonito
, “
An equivalent source technique for calculating the sound field inside an enclosure containing scattering objects
,”
J. Acoust. Soc. Am.
104
,
1221
1231
(
1998
).
34.
B.
Lam
,
S. J.
Elliott
,
J.
Cheer
, and
W.
Gan
, “
Regularisation of the equivalent source method for robust numerical modelling of acoustic scattering
,” in
Proceedings of Inter-Noise
(
2015
).
35.
Y. J. R.
Gounot
and
R. E.
Musafir
, “
Simulation of scattered fields: Some guidelines for the equivalent source method
,”
J. Sound Vib.
330
,
3698
3709
(
2011
).
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